Kala4,OsS1

• Symbol: Kala4,OsS1
• MSU: LOC_Os04g47059
• RAPdb: Os04g0557500

• The Birth of a Black Rice Gene and Its Local Spread by Introgression., 2015, Plant Cell.
• The C-S-A gene system regulates hull pigmentation and reveals evolution of anthocyanin biosynthesis pathway in rice., 2018, J Exp Bot.

Genbank accession number

• Here, we show that the black grain trait originated from ectopic expression of the Kala4 bHLH gene due to rearrangement in the promoter region

• Kala4~OsS1, Rc~qSD7-1~qPC7, The Birth of a Black Rice Gene and Its Local Spread by Introgression., Both the Rc and Kala4 genes activate upstream flavonol biosynthesis genes, such as chalcone synthase and dihydroflavonol-4-reductase, and downstream genes, such as leucoanthocyanidin reductase and leucoanthocyanidin dioxygenase, to produce the respective specific pigments